• Cellular and molecular mechanisms of morphogenesis in sponges (Porifera)

• Structural and functional characteristics of the biota of the White Sea rock pools

• Sprozoans (Apicomplexa, Sporozoa) and their hyperparasites

• Population investigations of marine benthos

• Three-spined stickleback as a key component of the White Sea ecosystem

• Spatio-temporal organization of microbenthic communities in the intertidal and upper subtidal zone of the White Sea

• Morphology and phylogeny of Turbellaria

• Algal ecophysiology and biochemistry